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REVIEW OF GENERA EVARCHA AND NIGORELLA, WITH COMMENTS ON EMERTONIUS, PADILOTHORAX, STAGETILLUS, AND DESCRIPTION OF FIVE NEW GENERA AND TWO NEW SPECIES (ARANEAE: SALTICIDAE)

Jerzy Prószyński

Abstract


The publication contains review of the genera of jumping spiders: Emertonius, Evarcha, Nigorella, Padilothorax, Stagetillus, and delimits five more new genera, provides graphic documentation for species considered recognizable and adds description of four new species. The taxonomic procedures are carried out according to methodology of "pragmatic classification", which stresses importance of graphic diagnostic characters presented in a comparative way (see also review  atMethodological postulates” subchapter). Some new procedures are introduced, testing their acceptability.

The following new genera are delimited and described in this paper: Evarcha Simon, 1889 s. s. (a part of Evarcha s. l.), Evacin Prószyński, 2017 gen. n., Evalba Prószyński, 2017 gen. n., Evaneg Prószyński, 2017 gen. n., Evawes Prószyński, 2017 gen. n., Padillothorus gen. n. Genus Emertonius Peckham & Peckham, 1892, misinterpreted twice by the WSC Editors, is reinstated again, with full diagnostic documentation repeated.

The following new species are described in this paper - Emertonius koomeni sp. n., Evacin besar sp. n., Evaneg aegiptiaca Prószyński, 2017 sp. n ."Emertonius" palawanensis sp. n.

New synonyms documented.

Evarcha acuta Wesolowska, 2006 = Evacin acuta (Wesolowska, 2006), comb. n., Evarcha bulbosa Zabka, 1985 = Evacin bulbosa (Zabka, 1985), comb. n., Evarcha cancellata Simon, 1902 = Evacin cancellata (Simon, 1902), comb. n., Evarcha flagellaris Haddad & Wesolowska, 2011= Evacin flagellaris (Haddad & Wesolowska, 2011), Evarcha flavocincta (C. L. Koch, 1846) = Evacin flavocincta (C. L. Koch, 1846) comb. n., Evarcha heteropogon Simon, 1903 = Evacin heteropogon Simon, 1903, comb. n., Evarcha infrastriata (Keyserling, 1881) = Evacin infrastriata (Keyserling, 1881), comb. n., Evarcha karas Wesolowska, 2011 = Evarcha karas (Wesolowska, 2011), comb. n., Evarcha kirghisica Rakov, 1997 = Evacin kirghisica (Rakov, 1997), comb. n., Evarcha kochi Simon, 1902 = Evacin kochi Simon, 1902 comb. n., Evarcha nigrifrons (Koch C.L., 1846) = Evacin nigrifrons (Koch C.L., 1846), comb. n., Evarcha optabilis Fox, 1937 = Evacin optabilis (Fox, 1937), comb. n., Evarcha pococki Zabka, 1985 = Evacin pococki (Zabka 1985), comb. n., Evarcha pulchella Thorell, 1895 = Evacin pulchella (Thorell, 1895), comb. n., Evarcha pseudopococki Peng X., Xie L. & Kim, 1993 = Evacin pseudopococki (Peng X., Xie L. & Kim, 1993), comb. n., Evarcha reiskindi Berry, Beatty, Proszynski, 1996 = Evacin reiskindi (Berry, Beatty, Proszynski, 1996), comb. n.,  Evarcha simoni[s] (Thorell, 1892)= Evacin simonis (Thorell, 1892), comb. n., Evarcha striolata Wesołowska & Haddad, 2009 = Evacin striolata (Wesołowska & Haddad, 2009), comb. n., Evarcha vitosa Próchniewicz, 1989 = Evacin vitosa (Próchniewicz, 1989), comb. n.

Evarcha albaria (L. Koch, 1878) = Evalba albaria (L. Koch, 1878), comb. n., Evarcha coreana Seo, 1988 = Evalba coreana (Seo, 1988), comb. n., Evarcha fasciata Seo, 1992 = Evalba fasciata (Seo, 1988), comb. n., Evarcha paralbaria Song & Chai, 1992 = Evalba paralbaria (Song & Chai, 1992), comb. n., Evarcha selenaria Suguro & Yahata, 2012 = Evalba selenaria (Suguro & Yahata, 2012), comb. n., Evarcha wulingensis Peng, Xie & Kim, 1993 = Evalba wulingensis (Peng, Xie & Kim, 1993), comb. n.

Evarcha armeniaca (Logunov, 1999) = Evaneg armeniaca (Logunov, 1999), comb. n., Evarcha darinurica Logunov, 2001 = Evaneg darinurica (Logunov, 2001) , comb. n., Evarcha negevensis (Proszynski, 2000) = Evaneg negevensis (Proszynski, 2000) comb. n., Evarcha nenilini Rakov, 1997= Evaneg nenilini (Rakov, 1997), comb. n., Evarcha nepos ( O. Pickard-Cambridge, 1872) = Evaneg nepos (O. Pickard-Cambridge, 1872), comb. n., Evarcha nigricans Dalmas, 1920 = Evaneg nigricans (Dalmas, 1920), comb. n.,

Evarcha pileckii Proszynski, 2000 = Evaneg pileckii (Proszynski, 2000), comb. n., Evarcha praeclara Prószyński & Wesolowska, in Prószyński, 2003 = Evaneg praeclara (Prószyński & Wesolowska, in Prószyński, 2003), comb. n., Evarcha seyun Wesolowska W., van Harten, 2007 = Evaneg seyun (Wesolowska W., van Harten, 2007), comb. n., Neaetha aegyptiaca Denis, 1947 female only = Evaneg aegyptiaca (Prószyński, 2017) ), comb. n., Neaetha aegyptiaca Denis, 1947 male only = Hyllus aegyptiacus (Denis, 1947), comb. n.

Evarcha arabica Wesolowska & van Harten, 2007 = Evawes arabica (Wesolowska & van Harten, 2007) comb.  n., Evarcha awashi Wesolowska & Tomasiewicz, 2008 = Evawes awashi (Wesolowska & Tomasiewicz, 2008) comb. n., Evarcha bakorensis Wesolowska & Russel-Smith 2011 = Evawes bakorensis (Wesolowska, Russel-Smith 2011) comb.n., Evarcha bicuspidata Peng & Li, 2003 = Evawes bicuspidata (Peng & Li, 2003) comb. n., Evarcha bihastata Wesolowska & Russell-Smith, 2000 = Evawes bihastata (Wesolowska & Russell-Smith, 2000), comb.n., Evarcha carbonaria (Lessert, 1927) = Evawes carbonaria (Lessert, 1927), comb.n., Evarcha chubbi Lessert, 1925 = Evawes chubbi (Lessert, 1925), comb. n., Evarcha culicivora Wesolowska & Jackson 2003 = Evawes culicivora (Wesolowska & Jackson 2003) comb. n., Evarcha denticulata Wesołowska & Haddad, 2013 = Evawes denticulata (Wesołowska & Haddad, 2013), comb.n., Evarcha elegans Wesolowska & Russel-Smith, 2000 = Evawes elegans (Wesolowska & Russel-Smith, 2000), comb. n., Evarcha ignea Wesołowska & Cumming 2008= Evawes ignea (Wesołowska & Cumming 2008), comb. n., Evarcha jucunda (Lucas, 1846) = Evawes jucunda (Lucas, 1846) comb. n., Evarcha longula (Thorell, 1881) = Evawes longula (Thorell, 1881) comb. n., Evarcha maculata Rollard & Wesolowska, 2002 = Evawes maculata (Rollard & Wesolowska, 2002), comb. n., Evarcha madagascariensis Proszynski, 1992 = Evawes madagascariensis (Proszynski, 1992), comb. n., Evarcha mirabilis Wesołowska & Haddad 2009 = Evawes mirabilis (Wesołowska & Haddad 2009), comb.n., Evarcha patagiata (O. Pickard-Cambridge, 1872) = Evawes patagiata (O. Pickard-Cambridge, 1872), comb.n., Evarcha picta Wesolowska & van Harten, 2007 = Evawes picta (Wesolowska & van Harten, 2007), comb. n., Evarcha pinguis Wesołowska &  Tomasiewicz, 2008 = Evawes pinguis (Wesołowska, Tomasiewicz, 2008), comb. n., Evarcha prosimilis Wesolowska & Cumming, 2008= Evawes prosimilis (Wesolowska & Cumming, 2008), comb. n., Evarcha zimbabwensis Wesolowska & Cumming, 2008 = Evawes zimbabwensis (Wesolowska & Cumming, 2008), comb. n.

Evarcha hirticeps (Song & Chai, 1992) = Nigorella hirticeps (Song & Chai, 1992), comb. n., Evarcha hunanensis Peng, Xie & Kim, 1993 - withdrawal from synonymy = Nigorella hunanensis (Peng, Xie & Kim, 1993), comb. n., Evarcha petrae Prószyński, 1992 = Nigorella petrae (Prószyński, 1992), comb. n., Evarcha sichuanensis Peng, Xie & Kim, 1993 = Nigorella sichuanensis (Peng, Xie & Kim, 1993), comb. n.

Hyllus fischeri Bösenberg & Strand, 1906 = Evacin fischeri (Bösenberg & Strand, 1906), comb. n. (separated from synonymy of Evarcha flavocincta).

Hasarius simonis Thorell, 1892 = Evacin simonis (Thorell, 1892), comb. n. (separated from synonymy of Evarcha flavocincta).

Myrmarachne aureonigra Edmunds, Prószyński, 2003) =  Myrmaplata aureonigra (Edmunds, Prószyński, 2003), comb. n.

Myrmarachne exasperans (Peckham & Peckham, 1892) = Emertonius exasperans Peckham & Peckham, 1892 (return to original combination),

"Myrmarachne" "exasperans": Wanless 1978: 235, f. 1A-F, not f. 2) (male from Palawan only) = "Emertonius" palawanensis sp. n.,

Myrmarachne grossa Edmunds & Prószyński, 2003 = Toxeus grossus (Edmunds & Prószyński, 2003) comb. n.

Neaetha aegyptiaca Denis, 1947 (in part - female) = Evaneg aegiptiaca Prószyński, 2017, sp. n., Neaetha aegyptiaca Denis, 1947 (in part - male) = Hyllus aegiptiacus (Denis, 1947) comb. n.

Stagetillus elegans (Reimoser, 1927) = Padillothorus elegans Reimoser, 1927, comb. n., reinstated original combination, Stagetillus semiostrinus (Simon, 1901) = Padillothorax semiostrinus Simon, 1901, comb. n., reinstated original combination, Stagetillus taprobanicus (Simon, 1902) = Padillothorax taprobanicus Simon, 1902, comb. n., reinstated original combination.

Viciria alba Peckham & Peckham, 1903 = Evacin alba (Peckham & Peckham, 1903), comb. n.

Correction of the following synonyms (rejecting biologically misleading changes by the WSC) documented in this paper.
Colyttus kerinci (Prószyński & Deeleman-Reinhold, 2012) = Donoessus kerinci Prószyński & Deeleman-Reinhold, 2012 (returned to the previous combination), Colyttus nigriceps (Simon, 1899) = Donoessus nigriceps (Simon, 1899) (returned to the previous combination), Colyttus striatus (Simon, 1902) = Donoessus striatus Simon, 1902 (returned to the previous combination),

Coryphasia heros (Bryant, 1943) = Dinattus heros Bryant, 1943 (returned to the previous combination), Jollas armatus (Bryant, 1943) = "Oningis" armatus Bryant, 1943 (returned to the previous combination), Jollas crassus (Bryant, 1943) = "Oningis" crassus Bryant, 1943 (returned to the previous combination), Laufeia daiqini (Prószyński & Deeleman-Reinhold, 2012) = Junxattus daiqini Prószyński & Deeleman-Reinhold, 2012 (returned to the previous combination), Laufeia eucola (Thorell, 1890) = Orcevia eucola Thorell, 1890 (returned to the previous combination), Laufeia keyserlingi (Thorell, 1890) = Orcevia keyserlingi Thorell, 1890 (returned to the previous combination), Laufeia kuloni (Prószyński & Deeleman-Reinhold, 2012) = Orcevia kuloni Prószyński & Deeleman-Reinhold, 2012, (returned to the previous combination,), Laufeia minuta (Prószyński, 1992) = Lechia minuta (Prószyński, 1992) (returned to the previous combination), Laufeia proszynskii Song, Gu & Chen, 1988= Orcevia proszynskii (Song, Gu & Chen, 1988) (returned to the previous combination), Laufeia squamata = Lechia squamata (returned to the previous combination)

Notice on pending revisions

Evaneg praeclara (Prószyński & Wesolowska in Prószyński, 2003) (= "Evarcha praeclara") is a composite species, pending taxonomic revision, "Myrmarachne" kilifi Wanless, 1978 - placement and diagnostic characters pending revision, "Myrmarachne" laurentina Bacelar, 1953 - placement and diagnostic characters pending revision.

Removal from Evarcha: Evarcha wenxianensis Tang & Yang, 1995 = Yaginumaella wenxianensis (Tang & Yang, 1995) (Fig, 15R). 

Keywords


Emertonius; Evarcha; Evacin; Evalba; Evaneg; Evawes; Padillothorus; Padillothorax; Stagetillus; Yaginumaella; Salticidae; new species; new genera; synonyms; pragmatic classification; molecular classification; chain of morphological similarities.

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