New data on the family Himantariidae Bollman, 1893 (Chilopoda: Geophilomorpha) from Kazakhstan

Polyporogaster porosa (Sseliwanoff, 1881) is new to the fauna of Kazakhstan; Bothriogaster signata (Kessler, 1874) is new to the fauna of the South Kazakhstan Region, while Stigmatogaster sp. is new to the East Kazakhstan Region. The distribution-map of all members of the family Himantariidae occurring in Kazakhstan and an identification key are provided.


Introduction
Two species of the family Himantariidae Bollman, 1893 have been recorded from Kazakhstan, viz. Bothriogaster signata (Kessler, 1874) and Polyporogaster schnitnikowi Lignau, 1929 (both Almaty Region), as well as specimens of the genus Stigmatogaster Latzel, 1880 (Tian-Shan Mts) (Lignau 1929a, b;Titova 1965Titova , 1978Dobroruka 1979); the considerable part of this large country has never been investigated. Based on new material from W, S and E Kazakhstan I provide new data on the fauna of these regions.

Materials and Methods
Present paper is based on the material collected in 2016-2018 (Fig. 1). Centipedes were taken by hand and preserved in 70% ethanol. The pictures have been taken using an Olympus DP74 digital camera attached to an Olympus SZX16 stereo microscope or Olympus XC50 camera attached to an Olympus BX51 microscope. The images were prepared with the help of Helicon Focus 6.7.1 and Paint.  Bothriogaster signata (Kessler, (Fig. 3) has numerous small denticles. Maxillary complex: telopodites of the first maxillae longer than the coxal projections; lappets of both coxosternite and telopodites absent. The second maxillae: coxosternite entire, its anterior margin slightly convex in the middle (arrow in Fig. 4). Forcipular coxosternite without denticles, chitin-lines reaching the condyles. Tarsungula, when closed, not reaching the anterior margin of the head (Fig. 2). Metasternites have pore-fields, large in anterior part of body and smaller in posterior part; two penultimate metasternites also have large pore-fields. Well-developed sternobothria accompanied by lateral gutters (Fig. 5) located mostly on 37-47 metasternites in male and on 40-53 metasternites in female. Ultimate metasternite wide, trapeziform with central longitudinal depression (Fig. 6). Coxal pores opening on ventral and dorsal sides in 2 groups (Figs 6, 7). Ultimate legs without pretarsus, swollen in male.
Remarks. The specimens studied agree well with the descriptions by Kessler (1874), Attems (1929), Lignau (1929a), Chalande and Ribaut (1909). This species is new to the fauna of the South Kazakhstan Region.
Remarks. This species is known from the original description only (Karatal River, vicinity of Balkhash Lake, Almaty Region) which is very succinct and provided with schematic figures of head (ventral view) and labrum. According to the original description this species is very close to the P. porosa; differs by shape of labral median arc only.

Genus Stigmatogaster Latzel, 1880
Stigmatogaster sp. Figs 1,[14][15][16][17][18][19][20] the curator of the myriapod collection in ZMMU, A.A. Schileyko (Moscow, Russia). I also wish to thank V.A. Krivokhatsky (Saint Petersburg, Russia) and G.Sh. Farzalieva (Perm, Russia) for the provision of material from the ZIN and PSU collections. I wish to thank M.M. Silantieva, Dean of the Institute of Biology and biotechnology of the ASU, as well as N.Yu. Speranskaya and M.Yu. Solomonova, staff members of the Department of Botany (all Barnaul) for their logistic help. Thanks go to the reviewers for corrections and critical commenting of the manuscript.