Description of the new genus Longarista Volynkin , gen . nov . , with a new species from northern Indochina ( Lepidoptera , Erebidae , Arctiinae )

The new genus Longarista Volynkin, gen. nov. is described for Miltochrista longaria Daniel, 1951. The new genus belongs to the Asura / Miltochrista generic complex and is related to Barsine Walker, 1854 and Miltochrista Hübner, [1819]. A new species, Longarista kareli Volynkin, sp. nov. is described from northern Indochina (Myanmar, Thailand, Laos and Vietnam). The new combination is established: Barsine convexa (Wileman, 1910) comb. nov. Adults, male and female genitalia of both Longarista species and representatives of the related genera are illustrated.


Introduction
The species M. longaria was described from southeastern China (Zhejiang) (Daniel, 1951) and long time was considered as a member of the genus Miltochrista (Fang, 1991;2000).Externally, this species is very similar to some species of the genus Barsine, but the study of its male and female genitalia demonstrated that it belongs not to Barsine or Miltochrista, but a separated, yet undescribed genus.In addition, during our studies of the genitalia of externally similar specimens from Myanmar, Thailand, Laos and Vietnam proved their belonging to another species, which is new to science.Descriptions of the new genus and the new species are given below.
The genitalia were dissected and mounted in euparal on glass slides.The photos of imagoes were taken using the camera Nikon D3100/AF-S Nikkor, 18-55 mm.The photos of genitalia where taken by the same camera attached to a microscope with an LM-scope adapter.All pictures were processed using the Adobe Photoshop CC 2018® software.
Longarista Volynkin, gen.nov.Type species: Miltochrista longaria Daniel, 1951.Etymology.The genus name is a combination of the name of the type species, longaria, and the generic name Miltochrista.Gender is feminine.
Diagnosis.Members of the new genus (Figs 1-9) resemble some species of the genus Barsine, especially females of the Vietnamese Barsine bachma bachma Volynkin & Černý, 2018 (Figs 13, 21, 28), males of the Chinese Barsine delineata (Walker, 1854) 20,27) and both sexes of Taiwanese Barsine convexa (Wileman, 1910) comb.nov.(Figs 14,15,22,29) as well.Nevertheless, species of Longarista can be easily distinguished from female specimens of B. bachma bachma by their broader forewing, presence of yellow suffusion and red hindwing cilia at outer and anal margin.Unlike B. delineata and B. convexa, species of Longarista have no discal spot of forewing.Despite the external similarity, the male genitalia of Longarista  differ clearly from those of  by the absence of a medial costal process, the presence of which is the main synapomorphy of the genus Barsine and its allies (Cyme Felder, 1861, Melanaema Butler, 1877, Barsura Volynkin, Dubatolov & Kishida, 2017, Nebulene Volynkin & Černý, 2018and Asura Walker, 1854).Compared to those of members of the genus Miltochrista (Figs 24,25), the male genitalia of Longarista have no distal membranous lobe of valva.In addition, the vesica of Longarista is armed with numerous small but robust cornuti, the feature bringing it closer to most Barsine (the vesica of Miltochrista has one or several large, thorn-or blade-like cornuti).The autapomorphic feature of Longarista is the structure of its juxta, which is swollen, heavily sclerotised and densely covered with small denticles, whereas other members of the Asura / Miltochrista generic complex have flattened juxta, which is moderately or weakly sclerotised and has no denticles.The female genitalia of the new genus (Fig. 26) are characterized by the presence of a swollen postvaginal plate, which is absent or flattened in  and other genera of the Asura / Miltochrista generic complex as well.1-9).Medium-sized moths; male antennae ciliate, female antennae filiform.Wings red with slight yellow suffusion.Forewing pattern dark grey or blackish, consists of two small subbasal spots, antemedial line angled in the cell, wavy medial line, curved and shortly dentate postmedial line and a series of longitudinal strokes of different length between the veins.Forewing cilia blackish.Hindwing without pattern, sometimes with slight dark grey suffusion along veins in the subterminal and terminal areas.Hindwing cilia blackish at apex and red at outer and anal margin.Male genitalia .Uncus robust, laterally flattened, curved subapically, pointed apically.Tuba analis broad, membranous; scaphium narrow and weakly sclerotised.Tegumen moderately sclerotised, shorter than valva, its distal half strongly angled dorsally; vinculum U-shaped, short; juxta bilobate, heavily sclerotised, strongly swollen, its dorsal surface densely dentate.Valva heavily sclerotized, lobe-like, with short broadly trigonal costal process directed distally and narrower thorn-like medial process directed distally-ventrally.Sacculus broad, weakly setose, without processes.Aedeagus elongated, narrow, slightly broadened and dentate distally.Vesica short, membranous, has long medial cluster of short but robust cornuti and several short diverticula with small short cornuti or granulation.Female genitalia (Fig. 26).Papillae anales trapezoidal.Apophyses thin; apophyses anteriores ca. 2 times shorter than apophyses posteriores.Ostium bursae broad.Postvaginal plate rounded, rugose, swollen dorsally.Ductus bursae elongated, dorso-ventrally flattened, heavily sclerotised.Corpus bursae ovoid, its inner surface with strong spinulose scobination.Appendix bursae small, globular, situated postero-laterally, with granulation laterally.

Other material
Diagnosis.Forewing length is 10-12 mm in males and 13 mm in females.Wingspan 20-25 mm in males is 26.5 mm in females.Males significantly vary in size and forewing width.The differences between L. longaria and L. kareli are listed in the diagnosis of the latter.
Diagnosis.Forewing length 10-12 mm in males.Wingspan is 20-25 mm.The new species (Figs 4-9) has no external differences from L. longaria (Figs 1-3) and can be distinguished by the genitalia structure only.Compared to those of L. longaria (Figs 16,17), the male genitalia of L. kareli (Figs 18,19) have the much narrower and less curved uncus, the slightly more elongated tegumen, the broader juxta with weaker dentation, the broader valva, and the broader aedeagus and vesica.Female is unknown.
Distribution.The new species is found in northern Myanmar (Kachin State), northern Thailand (Chiang Rai Province), northern Laos (Vientiane Province) and northern Vietnam (Hoa Binh Province).
Etymology.The species name is dedicated to Dr. Karel Černý, an expert in Asian Arctiinae and my friend.