A catalogue of the type specimens of Heteroptera ( Insecta ) housed at the Instituto Fundación Miguel Lillo ( Tucumán , Argentina )

This catalogue contains information about the type material of the suborder Heteroptera housed at the Entomological Collection of the Instituto Fundación Miguel Lillo (IFML-Tucumán, Argentina). We listed 60 holotypes and 453 paratypes belonging to 20 families, and three species and one subspecies that were not found in the collection but, according the original description, should be deposited in IFML. Finally, we listed 15 species that are labeled and coded as types but that are no part of the original type series. Nevertheless, they were included in this contribution because the associated information could be useful for future taxonomic research.


INTRODUCTION
With more than 40,000 described species, Heteroptera (Hemiptera), or true bugs, are part of the most successful radiation of nonholometabolous insects (Weirauch & Schuh, 2011).Heteropterans live on many different habitats: terrestrial, aquatic, intertidal and associated to other arthropods; and they include groups of phytophagous, predators and blood-sucking bugs.
Many species of heteropterans are known for their economical and/or medical concern, like species of the genus Triatoma Klug, vectors of the Chagas-Mazza disease (Schaefer & Panizzi, 2000).
The entomological collection of the Instituto Fundación Miguel Lillo (IFML -Tucumán, Argentina) is one of the most important in South America, including about 5 million specimens and type specimens of ca.5,500 species (Aranda et al., 2016); for a brief historical Table I review of the collection see Aranda et al. (2016).
Two previous works dealt with the type entomological collection at IFML, Hayward (1963) and Hayward (1967) that include true bugs families; both included a total of 67 species and subspecies of Heteroptera.In this contribution, we studied the type material of the suborder Heteroptera housed at the entomological collection of the IFML, increasing considerably the numbers given by Hayward and in many cases fixing misunderstandings and errors.

MATERIAL AND METHODS
The original combination of the type names is presented.The code THEM [number], provided by the IFML collection, refers to each type specimen of Hemiptera.The species are arranged alphabetically by genus following the classification of the suborder of Schuh & Slater (1995) and in the case of Lygaeoidea of Henry (1997).When the original description of the species has more information about the collecting data that is missing in the labels, it is given between brackets.
In the cases where more type material is deposited in other institutions or private collections, we mention them according to the data in the original description of the species.The acronyms used for the institutions are: Museo de La Plata, Argentina (MLP); Museo Argentino de Ciencias Naturales "Bernardino Rivadavia", Argentina (MACN); Instituto de Medicina Regional, Argentina Holotype: ♂ [Argentina], Jujuy, Abra Pampa, 3400 m, 5-6/III/1960, R. Golbach leg., THEM093.
Comments: Additional paratypes were deposited in MACN (three specimens), MLP (three specimens), and Bachmann's collection (three specimens).The author also mentioned four nymphs at IFML that were not found.
Comments: The male holotype and four paratypes were deposited in MACN, additional paratypes in MLP (three specimens), and Bachmann's collection (six specimens).Abdomen and genitalia of each male specimen glued to a separate cardboard.
Comments: Bachmann (1960) stated that the paratypes deposited in IFML are from the same locality than the holotype and several paratypes: "Río Negro, lagoons between Nahuel Huapi and Moreno lakes".The male holotype and 15 paratypes were deposited in MACN, additional paratypes in MLP (five, specimens), and Bachmann's collection (eight specimens).Comments: The male holotype and three paratypes were deposited in MACN, additional paratypes in MLP (two specimens), UKANS (two specimens), and Bachmann's collection (three specimens).Comments: The male holotype and ten paratypes were deposited in MACN, additional paratypes in MLP (three specimens), and Bachmann's collection (five specimens).According to the original description all specimens were collected by the same person.
Comments: The male holotype and additional nine paratypes were deposited in MACN, and one paratype in Bachmann's collection.One female paratype from "Tucumán, cerro San Javier" is missing in IFML.
Comments: We found some disagreements between the publication and the type material in IFML.According to the publication there are six female paratypes from "La Hoyada, Tucumán" and one male; and three females and two males from "Jujuy, Arroyo Los Berros".Additional paratypes were deposited in MLP (two specimens) and MACN (two specimens).Paratype: 1♀, [Argentina], Misiones, Loreto, THEM298.

CIMICOMORPHA
Comments: These specimens were originally deposited in IMR, then transferred to IFML.Additional paratypes were deposited in MACN (one specimen), MNRJ (one specimen), MCZ (one specimen), and Wygodzinsky's collection (three specimens).deposited in IMR, then transferred to IFML.Comments: This specimen was originally deposited in IMR, then transferred to IFML.The male holotype was deposited in CTIOC, and three additional paratypes in Wygodzinsky's collection.
Comments: These specimens were originally deposited in IMR, then transferred to IFML.Additional paratypes were deposited in MACN (three specimens), MLP (one specimen), and Wygodzinsky's collection (two specimens).
Comments: These specimens were originally deposited in IMR, then transferred to IFML.Comments: This specimen was originally deposited in IMR, then transferred to IFML.The male holotype was deposited in CEIOC, and two additional paratypes in Wygodzinsky's collection.The date in the original publication is 26-II-1947.

Stenolemus huali
Comments: This specimen was originally deposited in IMR, then transferred to IFML.
Comments: The male holotype was deposited in MACN; this paratype was originally deposited in IMR.
Comments: The male holotype and five paratypes were deposited in MEPRA, additional paratypes in CTIOC (two specimens), Prosen's (four specimens), Martinez's (four specimens), Pirán's (two specimens), Forster's, and Francisco Walz's collections (several paratypes without indication of the exact quantity).The specimen deposited in IFML is probably a paratype belonging to one of these private collections.

Phymata delpontei
Comments: The male holotype was deposited in Facultad de Agronomia y Veterinaria, Universidad de Buenos Aires; additional paratypes in CUIC (one specimen), ISM (one specimen), Elkins' (one specimen), and Lizer & Trelles' collections (six specimens).Comments: According to the original description two female paratypes from "Misiones, Iguazu" are deposited in Kormilev's collection and in ISM but none of them are indicated from "Alto Parana" and with a date.The holotype and three additional paratypes were deposited in Kormilev's collection, and one paratype in HNHM.Holotype: ♂, [Argentina], Sumampa, S.
Comments: The male holotype was deposited in MACN; the author did not mention the place where additional paratypes were deposited.The specimen labelled as THEM362 is not mentioned in the list of studied material.Comments: The male holotype was deposited in ISM.The author did not mention the place where the additional seven paratypes were deposited.Comments: These specimens were originally deposited in IMR, then transferred to IFML.Additional paratypes were deposited in USNM (one specimen), CTIOC (one specimen), Instituto de Higiene de Lima (Perú) (16 specimens), and Wygodzinsky's collection (two specimens).In the original publication three different localities near "Rio Marañon" are mentioned for the holotype and two paratypes, one of each corresponds to the one mentioned above (Jaen, 750 m, Yamón, 1100 m; and Lonya Grande, 1300 m).
Comments: According to Wygodzinsky (1948) the specimens are from the Northern forest of Perú and were reared in the laboratory.Four male paratypes that were originally deposited at IMR are missing.

MIROIDEA
Comments: One paratype was deposited in Carvalho's collection.
Comments: According to the original description the male holotype and one female paratype from "Santiago del Estero" should be deposited in the IFML collection.There is no indication of the depository of the other paratypes.However, if we assume that all of them should be in IFML: four ♂ and two ♀ paratypes from Santiago del Estero; one ♂ and one ♀ from Río Negro; two ♂ from La Rioja; two ♂ and three ♀ from Catamarca; two ♀ from Salta; and two ♀ from Córdoba are absent in the collection.
Comments: The paratypes were deposited in Carvalho's and Carpintero's collection.

ORTHOTYLINAE
Comments: According to the original description the male holotype should be deposited in IFML coded as AMNH_PBI 00195153 but it was not found in the collection; additional paratypes in AMNH (62 specimens) and USNM (83 specimens).
In the original description the female from "Jujuy" is referred from "Ceno Pezales"; there is one nymph instead of one male among the specimens from "Tartagal"; one female from "San Pedro de Colalao" collected by Monrós is missing; the material from "Burruyacu" is mentioned as collected in "20-I-948" and the material from "Lacavera" "22-11-51"; the specimens from "San Pedro de Colalao" collected by Willink are listed as deposited in IFML, MACN and Kormilev's collection.Finally, the female without data identified as THEM010 is not mentioned in the publication.Comments: The male holotype was deposited in the MNRJ, additional four paratypes in the MZSP.According to the original publication there should be four paratypes, a female from "28-I" missing in IFML.Comments: The male holotype and 72 paratypes were deposited in USNM, additional paratypes in AMNH (17 specimens) and Uhler's collection (81 specimens).The original publication mentions 69 paratypes, none from IFML.Comments: The male holotype and two paratypes were deposited in MACN, one additional paratype in MLP.According to the original description there should be two male and five female paratypes from Puerto Bemberg, Misiones in IFML.The male specimen has the hemelytra glued to a separate cardboard.Paratypes: 1♀, Perú, Tarma, 1-XII- [19]42, 3000 mts., W. Weirauch col., THEM132; 1♀, same data, THEM133.

COLOBATHRISTIDAE
Comments: The male holotype and two paratypes were deposited in AMNH, additional paratypes in MNHN (one specimen), and UNAM (two specimens).Comments: According to Brailovsky (1978), the male holotype and four paratypes were deposited in CAS; three paratypes in UNAM, and three in IFML.We found some disagreements between the publication and the paratypes examined in IFML, there is only one paratype from Río Luria, it is a male specimen and was deposited in UNAM.He also mentioned only male paratype/s from Huanuco deposited in IFML, but he did not mention the number of specimens, on the other hand the specimens THEM135 and THEM136 are from "Huanuco" and agree with the label data but are females.Comments: The male holotype and four paratypes were deposited in Investigaciones Entomológicas, Universidad de Chile, one paratype in MACN, and four paratypes in Kormilev's collection; probably one of these paratypes is the male deposited in IFML.Comments: The male holotype and one paratype were deposited in UNAM, additional paratypes in MNHN (two specimens) and USNM (one specimen).The original publication stated that two males and two females form "Ecuador: Provincia Tungurahua: Banios (1800 mts.), 4/5-IV-58, W. Weyrauch" are deposited in IFML and UNAM.Paratype: 1♀, Bolivia, Santa Cruz, Chiquitos, Santiago, 8-13/II/1958, Monrós col., Thaumastaneis sp.Det.M.V. Ahmat 1982, THEM123.Comments: The male holotype was deposited in MACN.Comments: Female holotype and two paratypes were deposited in AMNH, additional paratypes in BMNH (three specimens) and UNAM (five specimens).The original description mentioned only one female from "Alto del Tio" deposited in IFML.