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Cryptantha nevadensis var. rigida elevated to species with a new name, C. juniperensis



A replacement name, Cryptantha juniperensis, nomen et status novum, is proposed for Cryptantha nevadensis A.Nelson & P.B.Kennedy var. rigida I.M.Johnston. A new name was necessary because the name C. rigida was unavailable, due to its previous use for the South American Cryptantha rigida (Phil.) Reiche. Cryptantha juniperensis is similar to what has been treated as Cryptantha nevadensis var. nevadensis, with which it shares appressed upper stem trichomes and a relatively long fruiting calyx, often with erect to reflexed apices. However, Cryptantha juniperensis differs significantly from Cryptantha nevadensis var. nevadensis by the nutlet shape, which is narrowly ovate to widely lance-ovate, narrowly acute, and tuberculate in the former but lance-ovate, strongly acuminate, and elongate-tuberculate to muricate in the latter. In addition, Cryptantha juniperensis is erect (vs. lax and sprawling in Cryptantha nevadensis var. nevadensis), has fruiting calyces between 4.5–7.5 mm (vs. 6–11 mm long), and has a corolla limb that ranges between 2–5 mm in diameter (vs. 1–2 mm in diameter). Cryptantha juniperensis, which has also been classified as Cryptantha intermedia (A.Gray) Greene var. rigida Brand, is similar to recognized varieties of C. intermedia in nutlet morphology and in growth habit but differs in having upper stems with trichomes mostly appressed (vs. mostly spreading), a smaller corolla size (ca. 2–5 mm vs. 3–11 mm), and cymules that are mostly paired (vs. cymules that are arranged in threes). Preliminary molecular studies also support the recognition of C. juniperensis as a taxon separate from both C. nevadensis and C. intermedia.


Boraginaceae, nomenclature, taxonomy, systematics, Eudicots

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Abrams, L. (1951) Illustrated flora of the Pacific States: Washington, Oregon, and California. Vol. III Geraniaceae to Scrophulariaceae. Stanford University Press, Stanford, California.

Baja Flora. (2016) The Flora of Baja California. Available from: (accessed 8 December 2016)

Brand, A. (1931) Borraginaceae-Borraginoideae-Cryptantheae. In: Engler, A. (Eds.) Das Pflanzenreich 97. Verlag von Wilhelm Engelmann, Leipzig, 22 pp.

Brandegee, T.S. (1906) Plants of California. Zoe 5: 227–230.

Chacón, J., Luebert, F., Hilger, H.H., Ovcinnikova, S., Selvi, F., Cecchi, L., Guilliams, C.M., Hasenstab-Lehman, K., Sutorý, K., Simpson, M.G. & Weigend, M. (2016) The borage family (Boraginaceae s.str.): A revised infrafamilial classification based on new phylogenetic evidence, with emphasis on the placement of some enigmatic genera. Taxon 65: 523–546.

Consortium of California Herbaria (2016) Data provided by the participants of the Consortium of California Herbaria. Available from: (accessed 8 December 2016)

Cronquist, A. (1984) Cryptantha nevadensis treatment. In: A. Cronquist, A. H. Holmgren, N. Holmgren, H., J. L. Reveal, P. K. Holmgren (Eds.) Intermountain flora: Vascular plants of the intermountain west, U.S.A. Volume 4. Subclass Asteridae (except Asteraceae) 4. Subclass Asteridae (except Asteraceae). New York Botanic Garden Press, Bronx, New York, pp. 262.

Hasenstab-Lehman, K.E. & Simpson, M.G. (2012) Cat’s eyes and popcorn flowers: Phylogenetic systematics of the genus Cryptantha s.l. (Boraginaceae). Systematic Botany 37: 738–757.

Heller, A.A. (1906) Botanical exploration in California. Muhlenbergia 2: 1–416.

Johnston, I.M. (1925) Studies in the Boraginaceae IV. The North American species of Cryptantha. Contributions from the Gray Herbarium of Harvard University 74: 1–114.

Kearney, T.H. & Peebles, R.H. (1960) Arizona Flora. Ed. 2nd, with supplement by John Thomas Howell, Elizabeth McClintock, and collaborators. University of California Press, Berkeley.

Kelley, R.B., Simpson, M.G. & Hasenstab, K.E. (2012) Cryptantha. In: Baldwin, B.G., Goldman, D.H., Keil, D.J., Patterson, R., Rosatti, T.J. & Wilken, D.H. (Eds.) The Jepson Manual: Vascular Plants of California. Second Edition. University of California Press, Berkeley, Los Angeles, London, pp. 455–468.

Kelley, R.B. & Simpson, M.G. (In prep) Cryptantha. In: (Eds.) Flora of North America North of Mexico 16+ vols. Flora of North America Editorial Committee, New York and Oxford.

Kelley, W.A. & Wilken, D. (1993) Cryptantha. In: Hickman , J.C. (Eds.) The Jepson Manual: Higher Plants of California. University of California Press, Berkeley, pp. 369–378.

Luebert, F., Cecchi, L., Frohlich, M.W., Gottschling, M., Guilliams, C.M., Hasenstab-Lehman, K.E., Hilger, H.H., Miller, J.S., Mittelbach, M., Nazaire, M., Nepi, M., Nocentini, D., Ober, D., Olmstead, R.G., Selvi, F., Simpson, M.G., Sutorý, K., Valdés, B., Walden, G.K. & Weigend, M. (2016) Familial classification of the Boraginales. Taxon 65: 502–522.

Mabry, M.E. (2015) Evaluating the monophyly and biogeography of Cryptantha (Boraginaceae). Master of Science. San Diego State University, San Diego, California, 59 pp.

Macbride, J.F. (1916) III. Certain Borraginaceae [sic], new or transferred. Proceedings of the American Academy of Arts and Sciences 51: 541–548.

Nelson, A. & Kennedy, P.B. (1906) New Plants from the Great Basin. Proceedings of the Biological Society of Washington 19: 155–158.

Rydberg, P.A. (1909) Studies on the Rocky Mountain flora. Bulletin of the Torrey Botanical Club 36: 675–698.

SEINet. (2016) Southwest Environmental Information Network. Available from: http// (accessed 8 December 2016)

Simpson, M.G. (2008, onwards) Cryptantha Taxonomy & Images. Available from: (accessed July 2015)

Simpson, M.G. & Hasenstab, K.E. (2009) Cryptantha of Southern California. Crossosoma 35: 1–59.

Thiers, B. (2015, continuously updated) Index Herbariorum: A global directory of public herbaria and associated staff. Available from: (accessed June 2015)


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